1973

Habilitation in Genetics, University of Cologne

8/73 - 6/74

Postdoctoral training with F.H. Ruddle, Yale University, USA

1971 - 1976

Junior scientist, Institute of Genetics, University of Cologne

1968 - 1971

Postdoctoral training with A.B. Pardee, Princeton University, USA

1968

Ph.D. in Biochemistry with F. Lynen, University of Munich

1964

Diploma in Chemistry, University of Munich

1960 – 1964

Study of Chemistry at the Universities of Kiel and Munich

1/2009 - 8/2017

Senior professor at the University of Bonn

1/99-3/99

Research stay with B. Sutor, Institute of Physiology, University of Munich

10/93-3/94

Research stay with R. Jaenisch, Whitehead Institute, MIT, USA

1991- 1992

Chairman of Dept. of Biology (Fachgruppe Biologie), University of Bonn

1986 - 2008

C4 Professor of Genetics, University of Bonn, Director of the Institute of Genetics

10/82 - 5/83

Research stay with R. Schimke, Stanford University, USA

1977 - 1986

Associate Professor with tenure (C3), Institute of Cell Biology, University of Essen

1976

Associate Professor, University of Cologne

Several DFG stipends for postdoctoral and sabbatical work

1992

Max Planck Research Prize

1994-2008

Member of the scientific council of the German Cancer Aid, Dr. Mildred Scheel Foundation

2000-2003

Member of the Advisory Board, Heinrich Pette Institute, University of Hamburg